Iran buries slain Revolutionary Guards colonel

STORY: State television showed crowds surrounding a truck carrying Khodai's casket, wrapped in Iran's flag and strewn with flowers.

"Iran's response to any threat or action will be harsh. But we will determine when and how it will be and in what circumstances. We will definitely take revenge on our enemies," Revolutionary Guards commander Hossein Salami told reporters.

Khodai was shot dead by two people on a motorcycle in Tehran on Sunday (May 22).

The semi-official ISNA news agency said members of an Israeli intelligence service network had been discovered and arrested by the Guards.

The Israeli Prime Minister's Office, which oversees intelligence agency Mossad, declined to comment on the events in Tehran.

Israeli media said Khodai headed a unit of the Quds Force - the Revolutionary Guards' overseas arm - planning attacks on Israelis abroad.

The killing came at a time of uncertainty over the revival of Iran's 2015 nuclear deal with world powers after months of stalled talks.